The term used for where gears mesh is called 'pitch line'. On a gear, you have 'pitch circle diameter', which is given by a simple formula (PCD = Number of teeth x Module). Clearly, this formula breaks down for a rack, which is a gear with an infinite number of teeth. On a rack, the pitch line is where the tooth and the space are of equal thickness.
...I was hoping I could just download manufacturer step files of rack an pinion and then have Fusion 360 position them meshed correctly...
I think you are expecting too much. STEP files are the geometry of the part. The pitch circles of gears are derived properties. Would you expect to download a solid model and for it to indicate the centre of gravity of the part? Would you expect to download a 3D CAD file of a metric bolt and for it to show the pitch diameter of the thread?
Ime if you know what you are going to use then the manufacturer's will probably have a better model for you than a generic